David Pecker, the former publisher of the National Enquirer, testified at Donald Trump's trial Tuesday that the tabloid completely manufactured a negative story in 2016 about the father of Sen. Ted Cruz, of Texas, who was then Trump's rival for the GOP presidential nomination.
The paper had published a photo allegedly showing Cruz's father, Rafael Cruz, with Lee Harvey Oswald handing out pro-Fidel Castro pamphlets in New Orleans in 1963, not long before Oswald assassinated President John F. Kennedy.
Trump repeatedly referred to the story on the campaign trail and in interviews.
“I mean, what was he doing — what was he doing with Lee Harvey Oswald shortly before the death? Before the shooting?” Trump said in an interview with Fox News in May 2016. “It’s horrible.”
Get top local stories in DFW delivered to you every morning. Sign up for NBC DFW's News Headlines newsletter.
Manhattan prosecutor Joshua Steinglass asked Pecker about the story's origins during the trial Tuesday in Manhattan. Pecker said that then-National Enquirer editor-in-chief Dylan Howard and the tabloid's research department got involved, and Pecker indicated that they faked the photo that was the foundation for the story.
“We mashed the photos and the different picture with Lee Harvey Oswald. And mashed the two together. And that’s how that story was prepared — created I would say," Pecker said on the witness stand.
Asked by Steinglass whether Cruz had gained popularity in the presidential race at the time, Pecker said, “I believe so.”
The revelation came up as the prosecution focused on negative articles that were published by the tabloid about Trump's Republican opponents at the time. Pecker explained that it was Michael Cohen, Trump's personal lawyer, who would orchestrate the planting of these stories. Pecker said Cohen would call and say they'd like his publication to run an article on a certain candidate, adding that Cohen would then send him a piece about Cruz, for example, and the National Enquirer "would embellish it from there."
Pecker suggested that Trump was directly involved in the process, too. He said that the negative stories about Trump's opponents were published as part of an arrangement that was struck in 2015 at a Trump Tower meeting that also included a directive to write positive stories about the real estate mogul.
Steinglass also entered into evidence National Enquirer headlines published during the 2016 race about Sen. Marco Rubio, R-Fla., who was also running for president. They suggested that he had a love child and had a connection to cocaine.
Asked why the tabloid ran stories about the senators and candidate Ben Carson, Pecker said, "After the Republican debates, and based on the success that some of the other candidates had, I would receive a call from Michael Cohen, and he would direct me and direct Dylan Howard which candidate and which direction we should go.”
When asked for his response Tuesday, Cruz told NBC News he's “not interested in revisiting ancient history.”
Rubio's office did not immediately return a request for comment.
When the story about Cruz's dad was published, the senator told reporters that Trump was a "pathological liar" after he promoted the story.
“He doesn’t know the difference between truth and lies," he said. "He lies practically every word that comes out of his mouth. And in a pattern that I think is straight out of a psychology textbook, his response is to accuse everyone else of lying.”
This story first appeared on NBCNews.com. More from NBC News: